Autumn paints the valleys of Denali National Park in Alaska, home to more than 650 species of flowering plants. Fall brings changing patterns of color and motion to much of the United States, which covers lands from the tropics to the Arctic.

Here a grizzly bear forages in Denali’s 6 million acres (2.4 million hectares), bulking up for its winter hibernation. Also dependent on the park’s fall grasses, mosses, and shrubs are migrating birds and bull moose, which gather in their cows for the mating season.